Funding Information: We would like to thank Hyung Don Ryoo, Andreas Bergmann, Iswar Hariharan, Patrick Emery, the Bloomington Drosophila Stock Center, FlyORF and the Developmental Studies Hybridoma Bank (DSHB) for fly stocks, plasmids and antibodies. We would like to thank Hyung Don Ryoo, Avi Ashkenazi and Brenda Schulman for comments on the manuscript. We thank Julia Domingos (Insta: cenasdajul) for the art wo...