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Inhibition of glutamine synthetase by phosphinothricin leads to transcriptome reprograming in root nodules of Medicago truncatula

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Resumo:Glutamine synthetase (GS) is a vital enzyme for the assimilation of ammonia into amino acids in higher plants. In legumes, GS plays a crucial role in the assimilation of the ammonium released by nitrogen-fixing bacteria in root nodules, constituting an important metabolic knob controlling the nitrogen (N) assimilatory pathways. To identify new regulators of nodule metabolism, we profiled the transcriptome of Medicago truncatula nodules impaired in N assimilation by specifically inhibiting GS activity using phosphinothricin (PPT). Global transcript expression of nodules collected before and after PPT addition (4, 8, and 24 h) was assessed using Affymetrix M. truncatula GeneChip arrays. Hundreds of genes were regulated at the three time points, illustrating the dramatic alterations in cell metabolism that are imposed on the nodules upon GS inhibition. The data indicate that GS inhibition triggers a fast plant defense response, induces premature nodule senescence, and promotes loss of root nodule identity. Consecutive metabolic changes were identified at the three time points analyzed. The results point to a fast repression of asparagine synthesis and of the glycolytic pathway and to the synthesis of glutamate via reactions alternative to the GS/GOGAT cycle. Several genes potentially involved in the molecular surveillance for internal organic N availability are identified and a number of transporters potentially important for nodule functioning are pinpointed. The data provided by this study contributes to the mapping of regulatory and metabolic networks involved in root nodule functioning and highlight candidate modulators for functional analysis.
Autores principais:Seabra, Ana R
Outros Autores:Pereira, Patrícia A; Becker, Jörg D; Carvalho, Helena G
Assunto:Aminobutyrates Ammonium Compounds Enzyme Inhibitors Gene Expression Profiling Gene Expression Regulation, Plant Glutamate-Ammonia Ligase Medicago truncatula Nitrogen Nitrogen Fixation Oligonucleotide Array Sequence Analysis Plant Leaves Plant Proteins Plants, Genetically Modified RNA, Plant Root Nodules, Plant Sinorhizobium meliloti Symbiosis Time Factors Transcriptome
Ano:2012
País:Portugal
Tipo de documento:artigo
Tipo de acesso:acesso restrito
Instituição associada:Repositório do GIMM – Gulbenkian Institute for Molecular Medicine
Idioma:inglês
Origem:ARCA
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author Seabra, Ana R
author2 Pereira, Patrícia A
Becker, Jörg D
Carvalho, Helena G
author2_role author
author
author
author_facet Seabra, Ana R
Pereira, Patrícia A
Becker, Jörg D
Carvalho, Helena G
author_role author
contributor_name_str_mv ARCA
country_str PT
creators_json_txt [{\"Person.name\":\"Seabra, Ana R\"},{\"Person.name\":\"Pereira, Patrícia A\"},{\"Person.name\":\"Becker, Jörg D\"},{\"Person.name\":\"Carvalho, Helena G\"}]
datacite.contributors.contributor.contributorName.fl_str_mv ARCA
datacite.creators.creator.creatorName.fl_str_mv Seabra, Ana R
Pereira, Patrícia A
Becker, Jörg D
Carvalho, Helena G
datacite.date.Accepted.fl_str_mv 2012-07-01T00:00:00Z
datacite.date.available.fl_str_mv 2016-10-19T16:20:15Z
datacite.date.embargoed.fl_str_mv 2016-10-19T16:20:15Z
datacite.rights.fl_str_mv http://purl.org/coar/access_right/c_16ec
datacite.subjects.subject.fl_str_mv Aminobutyrates
Ammonium Compounds
Enzyme Inhibitors
Gene Expression Profiling
Gene Expression Regulation, Plant
Glutamate-Ammonia Ligase
Medicago truncatula
Nitrogen
Nitrogen Fixation
Oligonucleotide Array Sequence Analysis
Plant Leaves
Plant Proteins
Plants, Genetically Modified
RNA, Plant
Root Nodules, Plant
Sinorhizobium meliloti
Symbiosis
Time Factors
Transcriptome
datacite.titles.title.fl_str_mv Inhibition of glutamine synthetase by phosphinothricin leads to transcriptome reprograming in root nodules of Medicago truncatula
dc.contributor.none.fl_str_mv ARCA
dc.creator.none.fl_str_mv Seabra, Ana R
Pereira, Patrícia A
Becker, Jörg D
Carvalho, Helena G
dc.date.Accepted.fl_str_mv 2012-07-01T00:00:00Z
dc.date.available.fl_str_mv 2016-10-19T16:20:15Z
dc.date.embargoed.fl_str_mv 2016-10-19T16:20:15Z
dc.format.none.fl_str_mv application/pdf
dc.identifier.none.fl_str_mv http://hdl.handle.net/10400.7/704
dc.language.none.fl_str_mv eng
dc.publisher.none.fl_str_mv American Phytopathological Society
dc.rights.none.fl_str_mv http://purl.org/coar/access_right/c_16ec
dc.subject.none.fl_str_mv Aminobutyrates
Ammonium Compounds
Enzyme Inhibitors
Gene Expression Profiling
Gene Expression Regulation, Plant
Glutamate-Ammonia Ligase
Medicago truncatula
Nitrogen
Nitrogen Fixation
Oligonucleotide Array Sequence Analysis
Plant Leaves
Plant Proteins
Plants, Genetically Modified
RNA, Plant
Root Nodules, Plant
Sinorhizobium meliloti
Symbiosis
Time Factors
Transcriptome
dc.title.fl_str_mv Inhibition of glutamine synthetase by phosphinothricin leads to transcriptome reprograming in root nodules of Medicago truncatula
dc.type.none.fl_str_mv http://purl.org/coar/resource_type/c_6501
description Glutamine synthetase (GS) is a vital enzyme for the assimilation of ammonia into amino acids in higher plants. In legumes, GS plays a crucial role in the assimilation of the ammonium released by nitrogen-fixing bacteria in root nodules, constituting an important metabolic knob controlling the nitrogen (N) assimilatory pathways. To identify new regulators of nodule metabolism, we profiled the transcriptome of Medicago truncatula nodules impaired in N assimilation by specifically inhibiting GS activity using phosphinothricin (PPT). Global transcript expression of nodules collected before and after PPT addition (4, 8, and 24 h) was assessed using Affymetrix M. truncatula GeneChip arrays. Hundreds of genes were regulated at the three time points, illustrating the dramatic alterations in cell metabolism that are imposed on the nodules upon GS inhibition. The data indicate that GS inhibition triggers a fast plant defense response, induces premature nodule senescence, and promotes loss of root nodule identity. Consecutive metabolic changes were identified at the three time points analyzed. The results point to a fast repression of asparagine synthesis and of the glycolytic pathway and to the synthesis of glutamate via reactions alternative to the GS/GOGAT cycle. Several genes potentially involved in the molecular surveillance for internal organic N availability are identified and a number of transporters potentially important for nodule functioning are pinpointed. The data provided by this study contributes to the mapping of regulatory and metabolic networks involved in root nodule functioning and highlight candidate modulators for functional analysis.
dirty 0
eu_rights_str_mv restrictedAccess
format article
fulltext.url.fl_str_mv https://arca.gimm.pt/bitstreams/fce63352-07a9-408f-9829-bd064a7cadc0/download
funding.funder.alternateName_str_mv FCT
funding.funder.identifier_str_mv http://doi.org/10.13039/501100001871
funding.funder.name_str_mv Fundação para a Ciência e a Tecnologia
funding.identifier_str_mv PTDC/AGR-GPL/70592/2006
funding.name_str_mv 3599-PPCDT
funding_str_mv PTDC/AGR-GPL/70592/2006
info:eu-repo/grantAgreement/FCT/3599-PPCDT/PTDC%2FAGR-GPL%2F70592%2F2006/PT
id arca_4e9fc2bd0e8e5a2c3d4e0c2a41f2f164
identifier.url.fl_str_mv http://hdl.handle.net/10400.7/704
instacron_str arca
institution Repositório do GIMM – Gulbenkian Institute for Molecular Medicine
instname_str Repositório do GIMM – Gulbenkian Institute for Molecular Medicine
language eng
network_acronym_str arca
network_name_str ARCA
oai_identifier_str oai:arca.gimm.pt:10400.7/704
organization_str_mv urn:organizationAcronym:arca
person_str_mv Seabra, Ana R
Pereira, Patrícia A
Becker, Jörg D
Carvalho, Helena G
publishDate 2012
publisher.none.fl_str_mv American Phytopathological Society
reponame_str ARCA
repository_id_str urn:repositoryAcronym:arca
service_str_mv urn:repositoryAcronym:arca
spelling engAmerican Phytopathological Societypt_PTGlutamine synthetase (GS) is a vital enzyme for the assimilation of ammonia into amino acids in higher plants. In legumes, GS plays a crucial role in the assimilation of the ammonium released by nitrogen-fixing bacteria in root nodules, constituting an important metabolic knob controlling the nitrogen (N) assimilatory pathways. To identify new regulators of nodule metabolism, we profiled the transcriptome of Medicago truncatula nodules impaired in N assimilation by specifically inhibiting GS activity using phosphinothricin (PPT). Global transcript expression of nodules collected before and after PPT addition (4, 8, and 24 h) was assessed using Affymetrix M. truncatula GeneChip arrays. Hundreds of genes were regulated at the three time points, illustrating the dramatic alterations in cell metabolism that are imposed on the nodules upon GS inhibition. The data indicate that GS inhibition triggers a fast plant defense response, induces premature nodule senescence, and promotes loss of root nodule identity. Consecutive metabolic changes were identified at the three time points analyzed. The results point to a fast repression of asparagine synthesis and of the glycolytic pathway and to the synthesis of glutamate via reactions alternative to the GS/GOGAT cycle. Several genes potentially involved in the molecular surveillance for internal organic N availability are identified and a number of transporters potentially important for nodule functioning are pinpointed. The data provided by this study contributes to the mapping of regulatory and metabolic networks involved in root nodule functioning and highlight candidate modulators for functional analysis.application/pdfpt_PTInhibition of glutamine synthetase by phosphinothricin leads to transcriptome reprograming in root nodules of Medicago truncatulaSeabra, Ana RPereira, Patrícia ABecker, Jörg DCarvalho, Helena GHostingInstitutionOrganizationalARCAe-mailmailto:arca.repository@gimm.ptarca.repository@gimm.ptDOIIsPartOf10.1094/MPMI-12-11-03222016-10-19T16:20:15Z2012-072012-07-01T00:00:00ZHandlehttp://hdl.handle.net/10400.7/704http://purl.org/coar/access_right/c_16ecrestricted accessAminobutyratesAmmonium CompoundsEnzyme InhibitorsGene Expression ProfilingGene Expression Regulation, PlantGlutamate-Ammonia LigaseMedicago truncatulaNitrogenNitrogen FixationOligonucleotide Array Sequence AnalysisPlant LeavesPlant ProteinsPlants, Genetically ModifiedRNA, PlantRoot Nodules, PlantSinorhizobium melilotiSymbiosisTime FactorsTranscriptome774000 bytesFundação para a Ciência e a TecnologiaExpression profiling studies to specify the symbiotic genetic programme of the model legume Medicago truncatulainfo:eu-repo/grantAgreement/FCT/3599-PPCDT/PTDC%2FAGR-GPL%2F70592%2F2006/PTPTDC/AGR-GPL/70592/20063599-PPCDTCrossref Funder IDhttp://doi.org/10.13039/501100001871literaturehttp://purl.org/coar/resource_type/c_6501journal articlehttp://purl.org/coar/access_right/c_16ecapplication/pdffulltexthttps://arca.gimm.pt/bitstreams/fce63352-07a9-408f-9829-bd064a7cadc0/downloadMolecular Plant-Microbe Interactions257976992
spellingShingle Inhibition of glutamine synthetase by phosphinothricin leads to transcriptome reprograming in root nodules of Medicago truncatula
Seabra, Ana R
Aminobutyrates
Ammonium Compounds
Enzyme Inhibitors
Gene Expression Profiling
Gene Expression Regulation, Plant
Glutamate-Ammonia Ligase
Medicago truncatula
Nitrogen
Nitrogen Fixation
Oligonucleotide Array Sequence Analysis
Plant Leaves
Plant Proteins
Plants, Genetically Modified
RNA, Plant
Root Nodules, Plant
Sinorhizobium meliloti
Symbiosis
Time Factors
Transcriptome
status SINGLETON
subject.fl_str_mv Aminobutyrates
Ammonium Compounds
Enzyme Inhibitors
Gene Expression Profiling
Gene Expression Regulation, Plant
Glutamate-Ammonia Ligase
Medicago truncatula
Nitrogen
Nitrogen Fixation
Oligonucleotide Array Sequence Analysis
Plant Leaves
Plant Proteins
Plants, Genetically Modified
RNA, Plant
Root Nodules, Plant
Sinorhizobium meliloti
Symbiosis
Time Factors
Transcriptome
title Inhibition of glutamine synthetase by phosphinothricin leads to transcriptome reprograming in root nodules of Medicago truncatula
title_full Inhibition of glutamine synthetase by phosphinothricin leads to transcriptome reprograming in root nodules of Medicago truncatula
title_fullStr Inhibition of glutamine synthetase by phosphinothricin leads to transcriptome reprograming in root nodules of Medicago truncatula
title_full_unstemmed Inhibition of glutamine synthetase by phosphinothricin leads to transcriptome reprograming in root nodules of Medicago truncatula
title_short Inhibition of glutamine synthetase by phosphinothricin leads to transcriptome reprograming in root nodules of Medicago truncatula
title_sort Inhibition of glutamine synthetase by phosphinothricin leads to transcriptome reprograming in root nodules of Medicago truncatula
topic Aminobutyrates
Ammonium Compounds
Enzyme Inhibitors
Gene Expression Profiling
Gene Expression Regulation, Plant
Glutamate-Ammonia Ligase
Medicago truncatula
Nitrogen
Nitrogen Fixation
Oligonucleotide Array Sequence Analysis
Plant Leaves
Plant Proteins
Plants, Genetically Modified
RNA, Plant
Root Nodules, Plant
Sinorhizobium meliloti
Symbiosis
Time Factors
Transcriptome
topic_facet Aminobutyrates
Ammonium Compounds
Enzyme Inhibitors
Gene Expression Profiling
Gene Expression Regulation, Plant
Glutamate-Ammonia Ligase
Medicago truncatula
Nitrogen
Nitrogen Fixation
Oligonucleotide Array Sequence Analysis
Plant Leaves
Plant Proteins
Plants, Genetically Modified
RNA, Plant
Root Nodules, Plant
Sinorhizobium meliloti
Symbiosis
Time Factors
Transcriptome
url http://hdl.handle.net/10400.7/704
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