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The Impact of YabG Mutations on Clostridioides difficile Spore Germination and Processing of Spore Substrates

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Resumo:YabG is a sporulation-specific protease that is conserved among sporulating bacteria. Clostridioides difficile YabG processes the cortex destined proteins preproSleC into proSleC and CspBA to CspB and CspA. YabG also affects synthesis of spore coat/exosporium proteins CotA and CdeM. In prior work that identified CspA as the co-germinant receptor, mutations in yabG were found which altered the co-germinants required to initiate spore germination. To understand how these mutations in the yabG locus contribute to C. difficile spore germination, we introduced these mutations into an isogenic background. Spores derived from C. difficile yabGC207A (a catalytically inactive allele), C. difficile yabGA46D, C. difficile yabGG37E, and C. difficile yabGP153L strains germinated in response to taurocholic acid alone. Recombinantly expressed and purified preproSleC incubated with E. coli lysate expressing wild type YabG resulted in the removal of the presequence from preproSleC. Interestingly, only YabGA46D showed any activity toward purified preproSleC. Mutation of the YabG processing site in preproSleC (R119A) led to YabG shifting its processing to R115 or R112. Finally, changes in yabG expression under the mutant promoters were analyzed using a SNAP-tag and revealed expression differences at early and late stages of sporulation. Overall, our results support and expand upon the hypothesis that YabG is important for germination and spore assembly and, upon mutation of the processing site, can shift where it cleaves substrates.
Autores principais:Osborne, Morgan S
Outros Autores:Brehm, Joshua N; Olivença, Carmen; Cochran, Alicia M; Serrano, Mónica; Henriques, Adriano O; Sorg, Joseph A
Assunto:Clostridioides difficile germination protease spore YabG Microbiology Molecular Biology
Ano:2024
País:Portugal
Tipo de documento:artigo
Tipo de acesso:acesso aberto
Instituição associada:Universidade Nova de Lisboa
Idioma:inglês
Origem:Repositório Institucional da UNL
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author Osborne, Morgan S
author2 Brehm, Joshua N
Olivença, Carmen
Cochran, Alicia M
Serrano, Mónica
Henriques, Adriano O
Sorg, Joseph A
author2_role author
author
author
author
author
author
author_facet Osborne, Morgan S
Brehm, Joshua N
Olivença, Carmen
Cochran, Alicia M
Serrano, Mónica
Henriques, Adriano O
Sorg, Joseph A
author_role author
contributor_name_str_mv Instituto de Tecnologia Química e Biológica António Xavier (ITQB)
Molecular, Structural and Cellular Microbiology (MOSTMICRO)
Wiley
RUN
country_str PT
creators_json_txt [{\"Person.name\":\"Osborne, Morgan S\"},{\"Person.name\":\"Brehm, Joshua N\"},{\"Person.name\":\"Olivença, Carmen\"},{\"Person.name\":\"Cochran, Alicia M\"},{\"Person.name\":\"Serrano, Mónica\"},{\"Person.name\":\"Henriques, Adriano O\"},{\"Person.name\":\"Sorg, Joseph A\"}]
datacite.contributors.contributor.contributorName.fl_str_mv Instituto de Tecnologia Química e Biológica António Xavier (ITQB)
Molecular, Structural and Cellular Microbiology (MOSTMICRO)
Wiley
RUN
datacite.creators.creator.creatorName.fl_str_mv Osborne, Morgan S
Brehm, Joshua N
Olivença, Carmen
Cochran, Alicia M
Serrano, Mónica
Henriques, Adriano O
Sorg, Joseph A
datacite.date.Accepted.fl_str_mv 2024-09-11T00:00:00Z
datacite.date.available.fl_str_mv 2025-05-07T21:19:13Z
datacite.date.embargoed.fl_str_mv 2025-05-07T21:19:13Z
datacite.rights.fl_str_mv http://purl.org/coar/access_right/c_abf2
datacite.subjects.subject.fl_str_mv Clostridioides difficile
germination
protease
spore
YabG
Microbiology
Molecular Biology
datacite.titles.title.fl_str_mv The Impact of YabG Mutations on Clostridioides difficile Spore Germination and Processing of Spore Substrates
dc.contributor.none.fl_str_mv Instituto de Tecnologia Química e Biológica António Xavier (ITQB)
Molecular, Structural and Cellular Microbiology (MOSTMICRO)
Wiley
RUN
dc.creator.none.fl_str_mv Osborne, Morgan S
Brehm, Joshua N
Olivença, Carmen
Cochran, Alicia M
Serrano, Mónica
Henriques, Adriano O
Sorg, Joseph A
dc.date.Accepted.fl_str_mv 2024-09-11T00:00:00Z
dc.date.available.fl_str_mv 2025-05-07T21:19:13Z
dc.date.embargoed.fl_str_mv 2025-05-07T21:19:13Z
dc.format.none.fl_str_mv application/pdf
dc.identifier.none.fl_str_mv http://hdl.handle.net/10362/182744
dc.language.none.fl_str_mv eng
dc.rights.none.fl_str_mv http://purl.org/coar/access_right/c_abf2
dc.subject.none.fl_str_mv Clostridioides difficile
germination
protease
spore
YabG
Microbiology
Molecular Biology
dc.title.fl_str_mv The Impact of YabG Mutations on Clostridioides difficile Spore Germination and Processing of Spore Substrates
dc.type.none.fl_str_mv http://purl.org/coar/resource_type/c_6501
description YabG is a sporulation-specific protease that is conserved among sporulating bacteria. Clostridioides difficile YabG processes the cortex destined proteins preproSleC into proSleC and CspBA to CspB and CspA. YabG also affects synthesis of spore coat/exosporium proteins CotA and CdeM. In prior work that identified CspA as the co-germinant receptor, mutations in yabG were found which altered the co-germinants required to initiate spore germination. To understand how these mutations in the yabG locus contribute to C. difficile spore germination, we introduced these mutations into an isogenic background. Spores derived from C. difficile yabGC207A (a catalytically inactive allele), C. difficile yabGA46D, C. difficile yabGG37E, and C. difficile yabGP153L strains germinated in response to taurocholic acid alone. Recombinantly expressed and purified preproSleC incubated with E. coli lysate expressing wild type YabG resulted in the removal of the presequence from preproSleC. Interestingly, only YabGA46D showed any activity toward purified preproSleC. Mutation of the YabG processing site in preproSleC (R119A) led to YabG shifting its processing to R115 or R112. Finally, changes in yabG expression under the mutant promoters were analyzed using a SNAP-tag and revealed expression differences at early and late stages of sporulation. Overall, our results support and expand upon the hypothesis that YabG is important for germination and spore assembly and, upon mutation of the processing site, can shift where it cleaves substrates.
dirty 0
eu_rights_str_mv openAccess
format article
fulltext.url.fl_str_mv https://run.unl.pt/bitstreams/10d3d51c-46bf-4a36-8b15-a4e259873993/download
id run_a2ed8dccbff24d769bbfa09c22a7cd03
identifier.url.fl_str_mv http://hdl.handle.net/10362/182744
inst_facet_str urn:organizationAcronym:unl{{{_:::_}}}Universidade Nova de Lisboa
instacron_str unl
institution Universidade Nova de Lisboa
instname_str Universidade Nova de Lisboa
language eng
network_acronym_str run
network_name_str Repositório Institucional da UNL
oai_identifier_str oai:run.unl.pt:10362/182744
organization_str_mv urn:organizationAcronym:unl
person_str_mv Osborne, Morgan S
Brehm, Joshua N
Olivença, Carmen
Cochran, Alicia M
Serrano, Mónica
Henriques, Adriano O
Sorg, Joseph A
publishDate 2024
repo_facet_str urn:repositoryAcronym:run{{{_:::_}}}Repositório Institucional da UNL
reponame_str Repositório Institucional da UNL
repository_id_str urn:repositoryAcronym:run
service_str_mv urn:repositoryAcronym:run
spelling engenYabG is a sporulation-specific protease that is conserved among sporulating bacteria. Clostridioides difficile YabG processes the cortex destined proteins preproSleC into proSleC and CspBA to CspB and CspA. YabG also affects synthesis of spore coat/exosporium proteins CotA and CdeM. In prior work that identified CspA as the co-germinant receptor, mutations in yabG were found which altered the co-germinants required to initiate spore germination. To understand how these mutations in the yabG locus contribute to C. difficile spore germination, we introduced these mutations into an isogenic background. Spores derived from C. difficile yabGC207A (a catalytically inactive allele), C. difficile yabGA46D, C. difficile yabGG37E, and C. difficile yabGP153L strains germinated in response to taurocholic acid alone. Recombinantly expressed and purified preproSleC incubated with E. coli lysate expressing wild type YabG resulted in the removal of the presequence from preproSleC. Interestingly, only YabGA46D showed any activity toward purified preproSleC. Mutation of the YabG processing site in preproSleC (R119A) led to YabG shifting its processing to R115 or R112. Finally, changes in yabG expression under the mutant promoters were analyzed using a SNAP-tag and revealed expression differences at early and late stages of sporulation. Overall, our results support and expand upon the hypothesis that YabG is important for germination and spore assembly and, upon mutation of the processing site, can shift where it cleaves substrates.application/pdfenThe Impact of YabG Mutations on Clostridioides difficile Spore Germination and Processing of Spore SubstratesOsborne, Morgan SBrehm, Joshua NOlivença, CarmenCochran, Alicia MSerrano, MónicaHenriques, Adriano OSorg, Joseph AInstituto de Tecnologia Química e Biológica António Xavier (ITQB)Molecular, Structural and Cellular Microbiology (MOSTMICRO)WileyHostingInstitutionOrganizationalRUNe-mailmailto:run@unl.ptrun@unl.ptISSNIsPartOf0950-382XURNIsPartOfPURE: 99908043URNIsPartOfPURE UUID: eac966ca-926d-407a-9fe6-c42cf5111933URNIsPartOfScopus: 85203535325DOIIsPartOf10.1111/mmi.153162025-05-07T21:19:13Z2024-09-112024-09-11T00:00:00ZHandlehttp://hdl.handle.net/10362/182744http://purl.org/coar/access_right/c_abf2open accessClostridioides difficilegerminationproteasesporeYabGMicrobiologyMolecular Biology16198653 bytesliteraturehttp://purl.org/coar/resource_type/c_6501journal articlehttp://purl.org/coar/access_right/c_abf2application/pdffulltexthttps://run.unl.pt/bitstreams/10d3d51c-46bf-4a36-8b15-a4e259873993/download
spellingShingle The Impact of YabG Mutations on Clostridioides difficile Spore Germination and Processing of Spore Substrates
Osborne, Morgan S
Clostridioides difficile
germination
protease
spore
YabG
Microbiology
Molecular Biology
status SINGLETON
subject.fl_str_mv Clostridioides difficile
germination
protease
spore
YabG
Microbiology
Molecular Biology
title The Impact of YabG Mutations on Clostridioides difficile Spore Germination and Processing of Spore Substrates
title_full The Impact of YabG Mutations on Clostridioides difficile Spore Germination and Processing of Spore Substrates
title_fullStr The Impact of YabG Mutations on Clostridioides difficile Spore Germination and Processing of Spore Substrates
title_full_unstemmed The Impact of YabG Mutations on Clostridioides difficile Spore Germination and Processing of Spore Substrates
title_short The Impact of YabG Mutations on Clostridioides difficile Spore Germination and Processing of Spore Substrates
title_sort The Impact of YabG Mutations on Clostridioides difficile Spore Germination and Processing of Spore Substrates
topic Clostridioides difficile
germination
protease
spore
YabG
Microbiology
Molecular Biology
topic_facet Clostridioides difficile
germination
protease
spore
YabG
Microbiology
Molecular Biology
url http://hdl.handle.net/10362/182744
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