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Phosphoinositide 3-Kinase-Regulated pericyte maturation governs vascular remodeling

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Resumo:Background: Pericytes regulate vessel stabilization and function, and their loss is associated with diseases such as diabetic retinopathy or cancer. Despite their physiological importance, pericyte function and molecular regulation during angiogenesis remain poorly understood. Methods: To decipher the transcriptomic programs of pericytes during angiogenesis, we crossed Pdgfrb(BAC)-CreERT2 mice into RiboTagflox/flox mice. Pericyte morphological changes were assessed in mural cell-specific R26-mTmG reporter mice, in which low doses of tamoxifen allowed labeling of single-cell pericytes at high resolution. To study the role of phosphoinositide 3-kinase (PI3K) signaling in pericyte biology during angiogenesis, we used genetic mouse models that allow selective inactivation of PI3Kα and PI3Kβ isoforms and their negative regulator phosphate and tensin homolog deleted on chromosome 10 (PTEN) in mural cells. Results: At the onset of angiogenesis, pericytes exhibit molecular traits of cell proliferation and activated PI3K signaling, whereas during vascular remodeling, pericytes upregulate genes involved in mature pericyte cell function, together with a remarkable decrease in PI3K signaling. Immature pericytes showed stellate shape and high proliferation, and mature pericytes were quiescent and elongated. Unexpectedly, we demonstrate that PI3Kβ, but not PI3Kα, regulates pericyte proliferation and maturation during vessel formation. Genetic PI3Kβ inactivation in pericytes triggered early pericyte maturation. Conversely, unleashing PI3K signaling by means of PTEN deletion delayed pericyte maturation. Pericyte maturation was necessary to undergo vessel remodeling during angiogenesis. Conclusions: Our results identify new molecular and morphological traits associated with pericyte maturation and uncover PI3Kβ activity as a checkpoint to ensure appropriate vessel formation. In turn, our results may open new therapeutic opportunities to regulate angiogenesis in pathological processes through the manipulation of pericyte PI3Kβ activity.
Autores principais:Figueiredo, Ana
Outros Autores:Villacampa, Pilar; Diéguez-Hurtado, Rodrigo; José Lozano, Juan; Kobialka, Piotr; Cortazar, Ana Rosa; Martinez-Romero, Anabel; Angulo-Urarte, Ana; Franco, Claudio; Claret, Marc; Aransay, Ana Maria; Adams, Ralf H.; Carracedo, Arkaitz; Graupera, Mariona
Assunto:Angiogenesis, physiological Pericytes Phosphoinositide 3-kinase Signal transduction
Ano:2020
País:Portugal
Tipo de documento:artigo
Tipo de acesso:acesso restrito
Instituição associada:Universidade de Lisboa
Idioma:inglês
Origem:Repositório da Universidade de Lisboa
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author Figueiredo, Ana
author2 Villacampa, Pilar
Diéguez-Hurtado, Rodrigo
José Lozano, Juan
Kobialka, Piotr
Cortazar, Ana Rosa
Martinez-Romero, Anabel
Angulo-Urarte, Ana
Franco, Claudio
Claret, Marc
Aransay, Ana Maria
Adams, Ralf H.
Carracedo, Arkaitz
Graupera, Mariona
author2_role author
author
author
author
author
author
author
author
author
author
author
author
author
author_facet Figueiredo, Ana
Villacampa, Pilar
Diéguez-Hurtado, Rodrigo
José Lozano, Juan
Kobialka, Piotr
Cortazar, Ana Rosa
Martinez-Romero, Anabel
Angulo-Urarte, Ana
Franco, Claudio
Claret, Marc
Aransay, Ana Maria
Adams, Ralf H.
Carracedo, Arkaitz
Graupera, Mariona
author_role author
contributor_name_str_mv Repositório Científico de Acesso Aberto da ULisboa
country_str PT
creators_json_txt [{\"Person.name\":\"Figueiredo, Ana\",\"Person.identifier.orcid\":\"0000-0003-3499-9871\"},{\"Person.name\":\"Villacampa, Pilar\"},{\"Person.name\":\"Diéguez-Hurtado, Rodrigo\"},{\"Person.name\":\"José Lozano, Juan\"},{\"Person.name\":\"Kobialka, Piotr\"},{\"Person.name\":\"Cortazar, Ana Rosa\"},{\"Person.name\":\"Martinez-Romero, Anabel\"},{\"Person.name\":\"Angulo-Urarte, Ana\"},{\"Person.name\":\"Franco, Claudio\",\"Person.identifier.orcid\":\"0000-0002-2861-3883\"},{\"Person.name\":\"Claret, Marc\"},{\"Person.name\":\"Aransay, Ana Maria\",\"Person.identifier.orcid\":\"0000-0002-8271-612X\"},{\"Person.name\":\"Adams, Ralf H.\"},{\"Person.name\":\"Carracedo, Arkaitz\"},{\"Person.name\":\"Graupera, Mariona\"}]
datacite.contributors.contributor.contributorName.fl_str_mv Repositório Científico de Acesso Aberto da ULisboa
datacite.creators.creator.creatorName.fl_str_mv Figueiredo, Ana
Villacampa, Pilar
Diéguez-Hurtado, Rodrigo
José Lozano, Juan
Kobialka, Piotr
Cortazar, Ana Rosa
Martinez-Romero, Anabel
Angulo-Urarte, Ana
Franco, Claudio
Claret, Marc
Aransay, Ana Maria
Adams, Ralf H.
Carracedo, Arkaitz
Graupera, Mariona
datacite.date.Accepted.fl_str_mv 2020-08-18T00:00:00Z
datacite.date.available.fl_str_mv 2021-07-23T11:14:13Z
datacite.date.embargoed.fl_str_mv 2021-07-23T11:14:13Z
datacite.rights.fl_str_mv http://purl.org/coar/access_right/c_16ec
datacite.subjects.subject.fl_str_mv Angiogenesis, physiological
Pericytes
Phosphoinositide 3-kinase
Signal transduction
datacite.titles.title.fl_str_mv Phosphoinositide 3-Kinase-Regulated pericyte maturation governs vascular remodeling
dc.contributor.none.fl_str_mv Repositório Científico de Acesso Aberto da ULisboa
dc.creator.none.fl_str_mv Figueiredo, Ana
Villacampa, Pilar
Diéguez-Hurtado, Rodrigo
José Lozano, Juan
Kobialka, Piotr
Cortazar, Ana Rosa
Martinez-Romero, Anabel
Angulo-Urarte, Ana
Franco, Claudio
Claret, Marc
Aransay, Ana Maria
Adams, Ralf H.
Carracedo, Arkaitz
Graupera, Mariona
dc.date.Accepted.fl_str_mv 2020-08-18T00:00:00Z
dc.date.available.fl_str_mv 2021-07-23T11:14:13Z
dc.date.embargoed.fl_str_mv 2021-07-23T11:14:13Z
dc.format.none.fl_str_mv application/pdf
dc.identifier.none.fl_str_mv http://hdl.handle.net/10451/49078
dc.language.none.fl_str_mv eng
dc.publisher.none.fl_str_mv American Heart Association, Inc.
dc.rights.none.fl_str_mv http://purl.org/coar/access_right/c_16ec
dc.subject.none.fl_str_mv Angiogenesis, physiological
Pericytes
Phosphoinositide 3-kinase
Signal transduction
dc.title.fl_str_mv Phosphoinositide 3-Kinase-Regulated pericyte maturation governs vascular remodeling
dc.type.none.fl_str_mv http://purl.org/coar/resource_type/c_6501
description Background: Pericytes regulate vessel stabilization and function, and their loss is associated with diseases such as diabetic retinopathy or cancer. Despite their physiological importance, pericyte function and molecular regulation during angiogenesis remain poorly understood. Methods: To decipher the transcriptomic programs of pericytes during angiogenesis, we crossed Pdgfrb(BAC)-CreERT2 mice into RiboTagflox/flox mice. Pericyte morphological changes were assessed in mural cell-specific R26-mTmG reporter mice, in which low doses of tamoxifen allowed labeling of single-cell pericytes at high resolution. To study the role of phosphoinositide 3-kinase (PI3K) signaling in pericyte biology during angiogenesis, we used genetic mouse models that allow selective inactivation of PI3Kα and PI3Kβ isoforms and their negative regulator phosphate and tensin homolog deleted on chromosome 10 (PTEN) in mural cells. Results: At the onset of angiogenesis, pericytes exhibit molecular traits of cell proliferation and activated PI3K signaling, whereas during vascular remodeling, pericytes upregulate genes involved in mature pericyte cell function, together with a remarkable decrease in PI3K signaling. Immature pericytes showed stellate shape and high proliferation, and mature pericytes were quiescent and elongated. Unexpectedly, we demonstrate that PI3Kβ, but not PI3Kα, regulates pericyte proliferation and maturation during vessel formation. Genetic PI3Kβ inactivation in pericytes triggered early pericyte maturation. Conversely, unleashing PI3K signaling by means of PTEN deletion delayed pericyte maturation. Pericyte maturation was necessary to undergo vessel remodeling during angiogenesis. Conclusions: Our results identify new molecular and morphological traits associated with pericyte maturation and uncover PI3Kβ activity as a checkpoint to ensure appropriate vessel formation. In turn, our results may open new therapeutic opportunities to regulate angiogenesis in pathological processes through the manipulation of pericyte PI3Kβ activity.
dirty 0
eu_rights_str_mv restrictedAccess
format article
fulltext.url.fl_str_mv https://repositorio.ulisboa.pt/bitstreams/c6fe407c-1172-44b2-9cfe-e960cc2e7ff1/download
funding.funder.alternateName_str_mv EC
funding.funder.identifier_str_mv http://doi.org/10.13039/501100008530
funding.funder.name_str_mv European Commission
funding.name_str_mv H2020
id ul_7d4e5cbb3a059e095df0e7ad9d3930d5
identifier.url.fl_str_mv http://hdl.handle.net/10451/49078
instacron_str ul
institution Universidade de Lisboa
instname_str Universidade de Lisboa
language eng
network_acronym_str ul
network_name_str Repositório da Universidade de Lisboa
oai_identifier_str oai:repositorio.ulisboa.pt:10451/49078
organization_str_mv urn:organizationAcronym:ul
person_str_mv Figueiredo, Ana
Figueiredo, Ana
https://www.ciencia-id.pt/861B-2032-C580
861B-2032-C580
http://orcid.org/0000-0003-3499-9871
0000-0003-3499-9871
Villacampa, Pilar
Diéguez-Hurtado, Rodrigo
José Lozano, Juan
Kobialka, Piotr
Cortazar, Ana Rosa
Martinez-Romero, Anabel
Angulo-Urarte, Ana
Franco, Claudio
Franco, Claudio
https://www.ciencia-id.pt/F012-B7D6-AE72
F012-B7D6-AE72
http://orcid.org/0000-0002-2861-3883
0000-0002-2861-3883
Claret, Marc
Aransay, Ana Maria
Aransay, Ana Maria
http://orcid.org/0000-0002-8271-612X
0000-0002-8271-612X
Adams, Ralf H.
Carracedo, Arkaitz
Graupera, Mariona
publishDate 2020
publisher.none.fl_str_mv American Heart Association, Inc.
reponame_str Repositório da Universidade de Lisboa
repository_id_str urn:repositoryAcronym:ul
service_str_mv urn:repositoryAcronym:ul
spelling engAmerican Heart Association, Inc.pt_PTBackground: Pericytes regulate vessel stabilization and function, and their loss is associated with diseases such as diabetic retinopathy or cancer. Despite their physiological importance, pericyte function and molecular regulation during angiogenesis remain poorly understood. Methods: To decipher the transcriptomic programs of pericytes during angiogenesis, we crossed Pdgfrb(BAC)-CreERT2 mice into RiboTagflox/flox mice. Pericyte morphological changes were assessed in mural cell-specific R26-mTmG reporter mice, in which low doses of tamoxifen allowed labeling of single-cell pericytes at high resolution. To study the role of phosphoinositide 3-kinase (PI3K) signaling in pericyte biology during angiogenesis, we used genetic mouse models that allow selective inactivation of PI3Kα and PI3Kβ isoforms and their negative regulator phosphate and tensin homolog deleted on chromosome 10 (PTEN) in mural cells. Results: At the onset of angiogenesis, pericytes exhibit molecular traits of cell proliferation and activated PI3K signaling, whereas during vascular remodeling, pericytes upregulate genes involved in mature pericyte cell function, together with a remarkable decrease in PI3K signaling. Immature pericytes showed stellate shape and high proliferation, and mature pericytes were quiescent and elongated. Unexpectedly, we demonstrate that PI3Kβ, but not PI3Kα, regulates pericyte proliferation and maturation during vessel formation. Genetic PI3Kβ inactivation in pericytes triggered early pericyte maturation. Conversely, unleashing PI3K signaling by means of PTEN deletion delayed pericyte maturation. Pericyte maturation was necessary to undergo vessel remodeling during angiogenesis. Conclusions: Our results identify new molecular and morphological traits associated with pericyte maturation and uncover PI3Kβ activity as a checkpoint to ensure appropriate vessel formation. In turn, our results may open new therapeutic opportunities to regulate angiogenesis in pathological processes through the manipulation of pericyte PI3Kβ activity.application/pdfpt_PTPhosphoinositide 3-Kinase-Regulated pericyte maturation governs vascular remodelingPersonalFigueiredo, AnaDSpacehttp://dspace.org/items/8dc70955-51ce-45fa-b173-b236eba21681DSpacehttp://dspace.org/items/8dc70955-51ce-45fa-b173-b236eba21681Martins Figueiredo FonsecaAna RaquelCiência IDhttps://www.ciencia-id.pt861B-2032-C580ORCIDhttp://orcid.org0000-0003-3499-9871Villacampa, PilarDiéguez-Hurtado, RodrigoJosé Lozano, JuanKobialka, PiotrCortazar, Ana RosaMartinez-Romero, AnabelAngulo-Urarte, AnaPersonalFranco, ClaudioDSpacehttp://dspace.org/items/633fb0a6-1f81-4d35-98ca-b4cfd838d81eDSpacehttp://dspace.org/items/633fb0a6-1f81-4d35-98ca-b4cfd838d81eFrancoClaudioCiência IDhttps://www.ciencia-id.ptF012-B7D6-AE72ORCIDhttp://orcid.org0000-0002-2861-3883Scopus Author IDhttps://www.scopus.com24280736600Claret, MarcPersonalAransay, Ana MariaDSpacehttp://dspace.org/items/9ae0b1a0-3de0-477e-a83f-6710dbb86feaDSpacehttp://dspace.org/items/9ae0b1a0-3de0-477e-a83f-6710dbb86feaAransayAna MariaORCIDhttp://orcid.org0000-0002-8271-612XResearcher IDhttps://www.researcherid.comF-8086-2011Adams, Ralf H.Carracedo, ArkaitzGraupera, MarionaHostingInstitutionOrganizationalRepositório Científico de Acesso Aberto da ULisboae-mailmailto:repositorio@reitoria.ulisboa.ptrepositorio@reitoria.ulisboa.ptISSNIsPartOf0009-7322DOIIsPartOf10.1161/CIRCULATIONAHA.119.0423542021-07-23T11:14:13Z2020-08-182020-08-18T00:00:00ZHandlehttp://hdl.handle.net/10451/49078http://purl.org/coar/access_right/c_16ecrestricted accessAngiogenesis, physiologicalPericytesPhosphoinositide 3-kinaseSignal transduction6309276 bytesEuropean CommissionDeciphering PI3K biology in health and diseaseH2020Crossref Funder IDhttp://doi.org/10.13039/501100008530literaturehttp://purl.org/coar/resource_type/c_6501journal articlehttp://purl.org/coar/access_right/c_16ecapplication/pdffulltexthttps://repositorio.ulisboa.pt/bitstreams/c6fe407c-1172-44b2-9cfe-e960cc2e7ff1/downloadCirculation1427688704
spellingShingle Phosphoinositide 3-Kinase-Regulated pericyte maturation governs vascular remodeling
Figueiredo, Ana
Angiogenesis, physiological
Pericytes
Phosphoinositide 3-kinase
Signal transduction
status SINGLETON
subject.fl_str_mv Angiogenesis, physiological
Pericytes
Phosphoinositide 3-kinase
Signal transduction
title Phosphoinositide 3-Kinase-Regulated pericyte maturation governs vascular remodeling
title_full Phosphoinositide 3-Kinase-Regulated pericyte maturation governs vascular remodeling
title_fullStr Phosphoinositide 3-Kinase-Regulated pericyte maturation governs vascular remodeling
title_full_unstemmed Phosphoinositide 3-Kinase-Regulated pericyte maturation governs vascular remodeling
title_short Phosphoinositide 3-Kinase-Regulated pericyte maturation governs vascular remodeling
title_sort Phosphoinositide 3-Kinase-Regulated pericyte maturation governs vascular remodeling
topic Angiogenesis, physiological
Pericytes
Phosphoinositide 3-kinase
Signal transduction
topic_facet Angiogenesis, physiological
Pericytes
Phosphoinositide 3-kinase
Signal transduction
url http://hdl.handle.net/10451/49078
visible 1