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Lignin composition and structure differs between xylem, phloem and phellem in Quercus suber

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Resumo:The composition and structure of lignin in different tissues—phellem (cork), phloem and xylem (wood)—of Quercus suber was studied. Whole cell walls and their respective isolated milled lignins were analyzed by pyrolysis coupled with gas chromatography/mass spectrometry (Py-GC/MS), two-dimensional nuclear magnetic resonance spectroscopy (2D-NMR) and derivatization followed by reductive cleavage (DFRC). Different tissues presented varied p-hydroxyphenyl:guaiacyl:syringyl (H:G:S) lignin compositions. Whereas lignin from cork has a G-rich lignin (H:G:S molar ratio 2:85:13), lignin from phloem presents more S-units (H:G:S molar ratio of 1:58:41) and lignin from xylem is slightly enriched in S-lignin (H:G:S molar ratio 1:45:55). These differences were reflected in the relative abundances of the different interunit linkages. Alkyl-aryl ethers (β–O–4′) were predominant, increasing from 68% in cork, to 71% in phloem and 77% in xylem, as consequence of the enrichment in S-lignin units. Cork lignin was enriched in condensed structures such as phenylcoumarans (β-5′, 20%), dibenzodioxocins (5–5′, 5%), as corresponds to a lignin enriched in G-units. In comparison, lignin from phloem and xylem presented lower levels of condensed linkages. The lignin from cork was highly acetylated at the γ-OH of the side-chain (48% lignin acetylation), predominantly over G-units; while the lignins from phloem and xylem were barely acetylated and this occurred mainly over S-units. These results are a first time overview of the lignin structure in xylem, phloem (generated by cambium), and in cork (generated by phellogen), in agreement with literature that reports that lignin biosynthesis is flexible and cell specific.
Autores principais:Lourenço, Ana
Outros Autores:Rencoret, Jorge; Chemetova, Catarina; Gominho, Jorge; Gutierrez, Ana; Rio, José C. del; Pereira, Helena
Assunto:cork DFRC milled lignin NMR phloem Py-GC/MS Quercus suber xylem
Ano:2016
País:Portugal
Tipo de documento:artigo
Tipo de acesso:acesso aberto
Instituição associada:Universidade de Lisboa
Idioma:inglês
Origem:Repositório da Universidade de Lisboa
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author Lourenço, Ana
author2 Rencoret, Jorge
Chemetova, Catarina
Gominho, Jorge
Gutierrez, Ana
Rio, José C. del
Pereira, Helena
author2_role author
author
author
author
author
author
author_facet Lourenço, Ana
Lourenço, Ana
Rencoret, Jorge
Chemetova, Catarina
Gominho, Jorge
Gutierrez, Ana
Rio, José C. del
Pereira, Helena
Rencoret, Jorge
Chemetova, Catarina
Gominho, Jorge
Gutierrez, Ana
Rio, José C. del
Pereira, Helena
author_role author
contributor_name_str_mv Repositório Científico de Acesso Aberto da ULisboa
country_str PT
creators_json_str [{\"Person.name\":\"Lourenço, Ana\"},{\"Person.name\":\"Rencoret, Jorge\"},{\"Person.name\":\"Chemetova, Catarina\"},{\"Person.name\":\"Gominho, Jorge\"},{\"Person.name\":\"Gutierrez, Ana\"},{\"Person.name\":\"Rio, José C. del\"},{\"Person.name\":\"Pereira, Helena\"}]
datacite.contributors.contributor.contributorName.fl_str_mv Repositório Científico de Acesso Aberto da ULisboa
datacite.creators.creator.creatorName.fl_str_mv Lourenço, Ana
Rencoret, Jorge
Chemetova, Catarina
Gominho, Jorge
Gutierrez, Ana
Rio, José C. del
Pereira, Helena
datacite.date.Accepted.fl_str_mv 2016-10-01T00:00:00Z
datacite.date.available.fl_str_mv 2017-06-09T10:25:41Z
datacite.date.embargoed.fl_str_mv 2017-06-09T10:25:41Z
datacite.rights.fl_str_mv http://purl.org/coar/access_right/c_abf2
datacite.subjects.subject.fl_str_mv cork
DFRC
milled lignin
NMR
phloem
Py-GC/MS
Quercus suber
xylem
datacite.titles.title.fl_str_mv Lignin composition and structure differs between xylem, phloem and phellem in Quercus suber
dc.contributor.none.fl_str_mv Repositório Científico de Acesso Aberto da ULisboa
dc.creator.none.fl_str_mv Lourenço, Ana
Rencoret, Jorge
Chemetova, Catarina
Gominho, Jorge
Gutierrez, Ana
Rio, José C. del
Pereira, Helena
dc.date.Accepted.fl_str_mv 2016-10-01T00:00:00Z
dc.date.available.fl_str_mv 2017-06-09T10:25:41Z
dc.date.embargoed.fl_str_mv 2017-06-09T10:25:41Z
dc.format.none.fl_str_mv application/pdf
dc.identifier.none.fl_str_mv http://hdl.handle.net/10400.5/13741
dc.language.none.fl_str_mv eng
dc.publisher.none.fl_str_mv Danièle Werk (ed.)
dc.rights.none.fl_str_mv http://purl.org/coar/access_right/c_abf2
dc.subject.none.fl_str_mv cork
DFRC
milled lignin
NMR
phloem
Py-GC/MS
Quercus suber
xylem
dc.title.fl_str_mv Lignin composition and structure differs between xylem, phloem and phellem in Quercus suber
dc.type.none.fl_str_mv http://purl.org/coar/resource_type/c_6501
description The composition and structure of lignin in different tissues—phellem (cork), phloem and xylem (wood)—of Quercus suber was studied. Whole cell walls and their respective isolated milled lignins were analyzed by pyrolysis coupled with gas chromatography/mass spectrometry (Py-GC/MS), two-dimensional nuclear magnetic resonance spectroscopy (2D-NMR) and derivatization followed by reductive cleavage (DFRC). Different tissues presented varied p-hydroxyphenyl:guaiacyl:syringyl (H:G:S) lignin compositions. Whereas lignin from cork has a G-rich lignin (H:G:S molar ratio 2:85:13), lignin from phloem presents more S-units (H:G:S molar ratio of 1:58:41) and lignin from xylem is slightly enriched in S-lignin (H:G:S molar ratio 1:45:55). These differences were reflected in the relative abundances of the different interunit linkages. Alkyl-aryl ethers (β–O–4′) were predominant, increasing from 68% in cork, to 71% in phloem and 77% in xylem, as consequence of the enrichment in S-lignin units. Cork lignin was enriched in condensed structures such as phenylcoumarans (β-5′, 20%), dibenzodioxocins (5–5′, 5%), as corresponds to a lignin enriched in G-units. In comparison, lignin from phloem and xylem presented lower levels of condensed linkages. The lignin from cork was highly acetylated at the γ-OH of the side-chain (48% lignin acetylation), predominantly over G-units; while the lignins from phloem and xylem were barely acetylated and this occurred mainly over S-units. These results are a first time overview of the lignin structure in xylem, phloem (generated by cambium), and in cork (generated by phellogen), in agreement with literature that reports that lignin biosynthesis is flexible and cell specific.
dirty 0
eu_rights_str_mv openAccess
format article
fulltext.url.fl_str_mv https://repositorio.ulisboa.pt/bitstreams/9b3e9c55-9390-4ec1-86f9-b6b2b1e288e6/download
id ul_be7e1c1f6b4502b30b8a8e5300a79047
identifier.url.fl_str_mv http://hdl.handle.net/10400.5/13741
instacron_str ul
institution Universidade de Lisboa
instname_str Universidade de Lisboa
language eng
network_acronym_str ul
network_name_str Repositório da Universidade de Lisboa
oai_identifier_str oai:repositorio.ulisboa.pt:10400.5/13741
organization_str_mv urn:organizationAcronym:ul
person_str_mv Lourenço, Ana
Rencoret, Jorge
Chemetova, Catarina
Gominho, Jorge
Gutierrez, Ana
Rio, José C. del
Pereira, Helena
publishDate 2016
publisher.none.fl_str_mv Danièle Werk (ed.)
reponame_str Repositório da Universidade de Lisboa
repository_id_str urn:repositoryAcronym:ul
service_str_mv urn:repositoryAcronym:ul
spelling engDanièle Werk (ed.)pt_PTThe composition and structure of lignin in different tissues—phellem (cork), phloem and xylem (wood)—of Quercus suber was studied. Whole cell walls and their respective isolated milled lignins were analyzed by pyrolysis coupled with gas chromatography/mass spectrometry (Py-GC/MS), two-dimensional nuclear magnetic resonance spectroscopy (2D-NMR) and derivatization followed by reductive cleavage (DFRC). Different tissues presented varied p-hydroxyphenyl:guaiacyl:syringyl (H:G:S) lignin compositions. Whereas lignin from cork has a G-rich lignin (H:G:S molar ratio 2:85:13), lignin from phloem presents more S-units (H:G:S molar ratio of 1:58:41) and lignin from xylem is slightly enriched in S-lignin (H:G:S molar ratio 1:45:55). These differences were reflected in the relative abundances of the different interunit linkages. Alkyl-aryl ethers (β–O–4′) were predominant, increasing from 68% in cork, to 71% in phloem and 77% in xylem, as consequence of the enrichment in S-lignin units. Cork lignin was enriched in condensed structures such as phenylcoumarans (β-5′, 20%), dibenzodioxocins (5–5′, 5%), as corresponds to a lignin enriched in G-units. In comparison, lignin from phloem and xylem presented lower levels of condensed linkages. The lignin from cork was highly acetylated at the γ-OH of the side-chain (48% lignin acetylation), predominantly over G-units; while the lignins from phloem and xylem were barely acetylated and this occurred mainly over S-units. These results are a first time overview of the lignin structure in xylem, phloem (generated by cambium), and in cork (generated by phellogen), in agreement with literature that reports that lignin biosynthesis is flexible and cell specific.application/pdfpt_PTLignin composition and structure differs between xylem, phloem and phellem in Quercus suberLourenço, AnaRencoret, JorgeChemetova, CatarinaGominho, JorgeGutierrez, AnaRio, José C. delPereira, HelenaHostingInstitutionOrganizationalRepositório Científico de Acesso Aberto da ULisboae-mailmailto:repositorio@reitoria.ulisboa.ptrepositorio@reitoria.ulisboa.ptDOIIsPartOf10.3389/fpls.2016.016122017-06-09T10:25:41Z2016-102016-10-01T00:00:00ZHandlehttp://hdl.handle.net/10400.5/13741http://purl.org/coar/access_right/c_abf2open accesscorkDFRCmilled ligninNMRphloemPy-GC/MSQuercus suberxylem8904135 bytesliteraturehttp://purl.org/coar/resource_type/c_6501journal articlehttp://purl.org/coar/access_right/c_abf2application/pdffulltexthttps://repositorio.ulisboa.pt/bitstreams/9b3e9c55-9390-4ec1-86f9-b6b2b1e288e6/downloadFrontiers in Plant Science
spellingShingle Lignin composition and structure differs between xylem, phloem and phellem in Quercus suber
Lignin composition and structure differs between xylem, phloem and phellem in Quercus suber
Lourenço, Ana
cork
DFRC
milled lignin
NMR
phloem
Py-GC/MS
Quercus suber
xylem
Lourenço, Ana
cork
DFRC
milled lignin
NMR
phloem
Py-GC/MS
Quercus suber
xylem
status SINGLETON
subject.fl_str_mv cork
DFRC
milled lignin
NMR
phloem
Py-GC/MS
Quercus suber
xylem
title Lignin composition and structure differs between xylem, phloem and phellem in Quercus suber
title_full Lignin composition and structure differs between xylem, phloem and phellem in Quercus suber
title_fullStr Lignin composition and structure differs between xylem, phloem and phellem in Quercus suber
Lignin composition and structure differs between xylem, phloem and phellem in Quercus suber
title_full_unstemmed Lignin composition and structure differs between xylem, phloem and phellem in Quercus suber
Lignin composition and structure differs between xylem, phloem and phellem in Quercus suber
title_short Lignin composition and structure differs between xylem, phloem and phellem in Quercus suber
title_sort Lignin composition and structure differs between xylem, phloem and phellem in Quercus suber
topic cork
DFRC
milled lignin
NMR
phloem
Py-GC/MS
Quercus suber
xylem
topic_facet cork
DFRC
milled lignin
NMR
phloem
Py-GC/MS
Quercus suber
xylem
url http://hdl.handle.net/10400.5/13741
visible 1