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Drought reduces tree growing season lenght but increases nitrogen resorption efficiency in a Mediterranean ecosystem

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Resumo:Mediterranean ecosystems are hotspots for climate change, as the highest impacts are forecasted for the Mediterranean region, mainly by more frequent and intense severe droughts. Plant phenology is a good indicator of species’ responses to climate change. In this study, we compared the spring phenology of cork oak trees (Quercus suber), an evergreen species, over 2 contrasting years, a mild year (2004) and a dry year (2005), which was the most severe drought since records exist. We evaluated the timing of occurrence, duration, and intensity of bud development, budburst, shoot elongation, trunk growth, and leaf senescence (phenophases) and assessed the nitrogen resorption efficiency from senescent to green leaves. The temperature was the main driver of budburst. Nevertheless, water had the main role of constraining all the other phenophases by strongly reducing the growing season length (48 %) and consequently the tree growth. Basal area increment was the most affected growth variable (36 %), although it occurred at a similar rate in the 2 years. Shoot elongation was also reduced (21 %), yet elongation occurred at a higher rate in the dry year compared to the mild year. Leaf senescence during the bulk period was higher in the dry year, in which leaves were shed at the same rate over a longer period. Nitrogen concentrations in green and senescent leaves were affected by drought and nitrogen resorption efficiency increased remarkably (C22 %). Our results highlight the importance of studying different phenological metrics to improve our understanding of the ecosystem’s responses to climate change. The faster dynamics observed in shoot elongation, while all other phenophases developed at the same rate, indicate that leaf area development is privileged in cork oak. Water availability was the main driver of spring growth in this Mediterranean ecosystem; however, growth may be affected by complex interplays between precipitation and temperature, such as higher temperatures during dry winters or heatwaves during spring, that are likely to result in water stress. Longer studies are needed to disentangle those interplays. Finally, a higher nitrogen resorption efficiency in response to drought appears to be an adaptive trait that mitigates the limitation in nitrogen uptake by the roots during drought and contributes to improving tree fitness in the short term but will probably exert a negative feedback on the nitrogen cycle in the long term, which might affect the ecosystem functioning under the forecasted droughts
Autores principais:Lobo Do Vale, Raquel
Outros Autores:Besson, Cathy Kurz; Caldeira, Maria; Chaves, Maria Manuela; Pereira, João Santos
Assunto:drought tree growing nitrogen Mediterranean ecosystem
Ano:2019
País:Portugal
Tipo de documento:artigo
Tipo de acesso:acesso aberto
Instituição associada:Universidade de Lisboa
Idioma:português
Origem:Repositório da Universidade de Lisboa
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author Lobo Do Vale, Raquel
author2 Besson, Cathy Kurz
Caldeira, Maria
Chaves, Maria Manuela
Pereira, João Santos
author2_role author
author
author
author
author_facet Lobo Do Vale, Raquel
Lobo Do Vale, Raquel
Besson, Cathy Kurz
Caldeira, Maria
Chaves, Maria Manuela
Pereira, João Santos
Besson, Cathy Kurz
Caldeira, Maria
Chaves, Maria Manuela
Pereira, João Santos
author_role author
contributor_name_str_mv Repositório Científico de Acesso Aberto da ULisboa
country_str PT
creators_json_str [{\"Person.name\":\"Lobo Do Vale, Raquel\",\"Person.identifier.orcid\":\"0000-0003-4566-5420\"},{\"Person.name\":\"Besson, Cathy Kurz\"},{\"Person.name\":\"Caldeira, Maria\",\"Person.identifier.orcid\":\"0000-0002-3586-8526\"},{\"Person.name\":\"Chaves, Maria Manuela\"},{\"Person.name\":\"Pereira, João Santos\"}]
datacite.contributors.contributor.contributorName.fl_str_mv Repositório Científico de Acesso Aberto da ULisboa
datacite.creators.creator.creatorName.fl_str_mv Lobo Do Vale, Raquel
Besson, Cathy Kurz
Caldeira, Maria
Chaves, Maria Manuela
Pereira, João Santos
datacite.date.Accepted.fl_str_mv 2019-01-01T00:00:00Z
datacite.date.available.fl_str_mv 2019-04-11T10:14:07Z
datacite.date.embargoed.fl_str_mv 2019-04-11T10:14:07Z
datacite.rights.fl_str_mv http://purl.org/coar/access_right/c_abf2
datacite.subjects.subject.fl_str_mv drought
tree growing
nitrogen
Mediterranean ecosystem
datacite.titles.title.fl_str_mv Drought reduces tree growing season lenght but increases nitrogen resorption efficiency in a Mediterranean ecosystem
dc.contributor.none.fl_str_mv Repositório Científico de Acesso Aberto da ULisboa
dc.creator.none.fl_str_mv Lobo Do Vale, Raquel
Besson, Cathy Kurz
Caldeira, Maria
Chaves, Maria Manuela
Pereira, João Santos
dc.date.Accepted.fl_str_mv 2019-01-01T00:00:00Z
dc.date.available.fl_str_mv 2019-04-11T10:14:07Z
dc.date.embargoed.fl_str_mv 2019-04-11T10:14:07Z
dc.format.none.fl_str_mv application/pdf
application/pdf
dc.identifier.none.fl_str_mv http://hdl.handle.net/10400.5/17726
dc.language.none.fl_str_mv por
dc.publisher.none.fl_str_mv European Geosciences Union
dc.rights.none.fl_str_mv http://purl.org/coar/access_right/c_abf2
dc.subject.none.fl_str_mv drought
tree growing
nitrogen
Mediterranean ecosystem
dc.title.fl_str_mv Drought reduces tree growing season lenght but increases nitrogen resorption efficiency in a Mediterranean ecosystem
dc.type.none.fl_str_mv http://purl.org/coar/resource_type/c_6501
description Mediterranean ecosystems are hotspots for climate change, as the highest impacts are forecasted for the Mediterranean region, mainly by more frequent and intense severe droughts. Plant phenology is a good indicator of species’ responses to climate change. In this study, we compared the spring phenology of cork oak trees (Quercus suber), an evergreen species, over 2 contrasting years, a mild year (2004) and a dry year (2005), which was the most severe drought since records exist. We evaluated the timing of occurrence, duration, and intensity of bud development, budburst, shoot elongation, trunk growth, and leaf senescence (phenophases) and assessed the nitrogen resorption efficiency from senescent to green leaves. The temperature was the main driver of budburst. Nevertheless, water had the main role of constraining all the other phenophases by strongly reducing the growing season length (48 %) and consequently the tree growth. Basal area increment was the most affected growth variable (36 %), although it occurred at a similar rate in the 2 years. Shoot elongation was also reduced (21 %), yet elongation occurred at a higher rate in the dry year compared to the mild year. Leaf senescence during the bulk period was higher in the dry year, in which leaves were shed at the same rate over a longer period. Nitrogen concentrations in green and senescent leaves were affected by drought and nitrogen resorption efficiency increased remarkably (C22 %). Our results highlight the importance of studying different phenological metrics to improve our understanding of the ecosystem’s responses to climate change. The faster dynamics observed in shoot elongation, while all other phenophases developed at the same rate, indicate that leaf area development is privileged in cork oak. Water availability was the main driver of spring growth in this Mediterranean ecosystem; however, growth may be affected by complex interplays between precipitation and temperature, such as higher temperatures during dry winters or heatwaves during spring, that are likely to result in water stress. Longer studies are needed to disentangle those interplays. Finally, a higher nitrogen resorption efficiency in response to drought appears to be an adaptive trait that mitigates the limitation in nitrogen uptake by the roots during drought and contributes to improving tree fitness in the short term but will probably exert a negative feedback on the nitrogen cycle in the long term, which might affect the ecosystem functioning under the forecasted droughts
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eu_rights_str_mv openAccess
format article
fulltext.url.fl_str_mv https://repositorio.ulisboa.pt/bitstreams/034e73a2-d05e-4a5c-8c10-3b02f2d5da70/download
id ul_fd41442802460a2b8d655ea925fc39bb
identifier.url.fl_str_mv http://hdl.handle.net/10400.5/17726
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institution Universidade de Lisboa
instname_str Universidade de Lisboa
language por
network_acronym_str ul
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oai_identifier_str oai:repositorio.ulisboa.pt:10400.5/17726
organization_str_mv urn:organizationAcronym:ul
person_str_mv Lobo Do Vale, Raquel
Lobo Do Vale, Raquel
https://www.ciencia-id.pt/2911-9D6C-E289
2911-9D6C-E289
http://orcid.org/0000-0003-4566-5420
0000-0003-4566-5420
Besson, Cathy Kurz
Caldeira, Maria
Caldeira, Maria
https://www.ciencia-id.pt/C816-E9F6-8272
C816-E9F6-8272
http://orcid.org/0000-0002-3586-8526
0000-0002-3586-8526
Chaves, Maria Manuela
Pereira, João Santos
publishDate 2019
publisher.none.fl_str_mv European Geosciences Union
reponame_str Repositório da Universidade de Lisboa
repository_id_str urn:repositoryAcronym:ul
service_str_mv urn:repositoryAcronym:ul
spelling porEuropean Geosciences Unionpt_PTMediterranean ecosystems are hotspots for climate change, as the highest impacts are forecasted for the Mediterranean region, mainly by more frequent and intense severe droughts. Plant phenology is a good indicator of species’ responses to climate change. In this study, we compared the spring phenology of cork oak trees (Quercus suber), an evergreen species, over 2 contrasting years, a mild year (2004) and a dry year (2005), which was the most severe drought since records exist. We evaluated the timing of occurrence, duration, and intensity of bud development, budburst, shoot elongation, trunk growth, and leaf senescence (phenophases) and assessed the nitrogen resorption efficiency from senescent to green leaves. The temperature was the main driver of budburst. Nevertheless, water had the main role of constraining all the other phenophases by strongly reducing the growing season length (48 %) and consequently the tree growth. Basal area increment was the most affected growth variable (36 %), although it occurred at a similar rate in the 2 years. Shoot elongation was also reduced (21 %), yet elongation occurred at a higher rate in the dry year compared to the mild year. Leaf senescence during the bulk period was higher in the dry year, in which leaves were shed at the same rate over a longer period. Nitrogen concentrations in green and senescent leaves were affected by drought and nitrogen resorption efficiency increased remarkably (C22 %). Our results highlight the importance of studying different phenological metrics to improve our understanding of the ecosystem’s responses to climate change. The faster dynamics observed in shoot elongation, while all other phenophases developed at the same rate, indicate that leaf area development is privileged in cork oak. Water availability was the main driver of spring growth in this Mediterranean ecosystem; however, growth may be affected by complex interplays between precipitation and temperature, such as higher temperatures during dry winters or heatwaves during spring, that are likely to result in water stress. Longer studies are needed to disentangle those interplays. Finally, a higher nitrogen resorption efficiency in response to drought appears to be an adaptive trait that mitigates the limitation in nitrogen uptake by the roots during drought and contributes to improving tree fitness in the short term but will probably exert a negative feedback on the nitrogen cycle in the long term, which might affect the ecosystem functioning under the forecasted droughtsapplication/pdfapplication/pdfpt_PTDrought reduces tree growing season lenght but increases nitrogen resorption efficiency in a Mediterranean ecosystemPersonalLobo Do Vale, RaquelDSpacehttp://dspace.org/items/16e8670c-9a61-4a5e-b5c7-c47023b566c3DSpacehttp://dspace.org/items/16e8670c-9a61-4a5e-b5c7-c47023b566c3Lobo do ValeRaquelCiência IDhttps://www.ciencia-id.pt2911-9D6C-E289ORCIDhttp://orcid.org0000-0003-4566-5420Researcher IDhttps://www.researcherid.comD-4570-2013Scopus Author IDhttps://www.scopus.com20436288000Besson, Cathy KurzPersonalCaldeira, MariaDSpacehttp://dspace.org/items/8d30abe1-834f-4610-8f7e-62c0add60582DSpacehttp://dspace.org/items/8d30abe1-834f-4610-8f7e-62c0add60582da Conceição Brálio de Brito CaldeiraMariaCiência IDhttps://www.ciencia-id.ptC816-E9F6-8272ORCIDhttp://orcid.org0000-0002-3586-8526Chaves, Maria ManuelaPereira, João SantosHostingInstitutionOrganizationalRepositório Científico de Acesso Aberto da ULisboae-mailmailto:repositorio@reitoria.ulisboa.ptrepositorio@reitoria.ulisboa.ptDOIIsPartOfhttps://doi.org/10.5194/bg-16-1265-20192019-04-11T10:14:07Z20192019-01-01T00:00:00ZHandlehttp://hdl.handle.net/10400.5/17726http://purl.org/coar/access_right/c_abf2open accessdroughttree growingnitrogenMediterranean ecosystem631447 bytes772479 bytesliteraturehttp://purl.org/coar/resource_type/c_6501journal articlehttp://purl.org/coar/access_right/c_abf2application/pdffulltexthttps://repositorio.ulisboa.pt/bitstreams/034e73a2-d05e-4a5c-8c10-3b02f2d5da70/downloadhttp://purl.org/coar/access_right/c_abf2application/pdffulltexthttps://repositorio.ulisboa.pt/bitstreams/54a5952e-3b1e-483c-af5c-487f5b0ad70b/downloadBiogeosciences
spellingShingle Drought reduces tree growing season lenght but increases nitrogen resorption efficiency in a Mediterranean ecosystem
Drought reduces tree growing season lenght but increases nitrogen resorption efficiency in a Mediterranean ecosystem
Lobo Do Vale, Raquel
drought
tree growing
nitrogen
Mediterranean ecosystem
Lobo Do Vale, Raquel
drought
tree growing
nitrogen
Mediterranean ecosystem
status SINGLETON
subject.fl_str_mv drought
tree growing
nitrogen
Mediterranean ecosystem
title Drought reduces tree growing season lenght but increases nitrogen resorption efficiency in a Mediterranean ecosystem
title_full Drought reduces tree growing season lenght but increases nitrogen resorption efficiency in a Mediterranean ecosystem
title_fullStr Drought reduces tree growing season lenght but increases nitrogen resorption efficiency in a Mediterranean ecosystem
Drought reduces tree growing season lenght but increases nitrogen resorption efficiency in a Mediterranean ecosystem
title_full_unstemmed Drought reduces tree growing season lenght but increases nitrogen resorption efficiency in a Mediterranean ecosystem
Drought reduces tree growing season lenght but increases nitrogen resorption efficiency in a Mediterranean ecosystem
title_short Drought reduces tree growing season lenght but increases nitrogen resorption efficiency in a Mediterranean ecosystem
title_sort Drought reduces tree growing season lenght but increases nitrogen resorption efficiency in a Mediterranean ecosystem
topic drought
tree growing
nitrogen
Mediterranean ecosystem
topic_facet drought
tree growing
nitrogen
Mediterranean ecosystem
url http://hdl.handle.net/10400.5/17726
visible 1